PMID:10102273 in vitro, by Akt, phosphosites identified by mutagenesis in 293T using phosphoFOXO1 antibodies PMID:18593906 in HEK293T, by PIM1s isoform, FOXO3 phosphorylated and inactivated. Phosphomutant of FOXO3 is not phosphorylated and the expression of a reporter gene is not inhibited.