(1) FZR1|pho@nucleus -> FZR1@nucleus
(2) FZR1@nucleus -> FZR1|pho@nucleus
(3) ANAPC1:ANAPC10:ANAPC11:ANAPC13:ANAPC2:ANAPC4:ANAPC5:ANAPC7:CDC16:CDC23:CDC26:CDC27@nucleus+FZR1@nucleus -> ANAPC1:ANAPC10:ANAPC11:ANAPC13:ANAPC2:ANAPC4:ANAPC5:ANAPC7:CDC16:CDC23:CDC26:CDC27:FZR1@nucleus